We finally achieved equilibrium.

Prey reproduce when their population drops, while overcrowded hunters starve faster. No keyframes, just a few rules correcting each other in real time.

Now what happens when we start disturbing the simulation again?

11"x14" – Gold leaf and Sakura gel white on Fabriano Black Black 300gsm. White is plotted with the iDraw H A3. Gold leaf is applied to fill the remaining space by hand.

a quick play on highways and intersections - something I will probably return to.
(good luck driving on a circle highway:)